This is a text which is used in my Genome 562 course. It was originally
written in the late 1970's, and many parts of it have not been updated since
about 1981. A major part of the new chapter on molecular population genetics
and most of a chapter on multiple linked loci, plus a chapter on polygenic
characters in natural populations, has been added more recently.
It was to be a book, but I became involved in work on phylogenies,
and it has been published as lecture notes ever since. A small amount of new
material has been added each time it was privately reprinted by me for use
in my class. It was originally on a magnetic-card storage system for
electric typewriters, and then migrated to the Runoff family of mathematical
text formatting systems, and finally to LaTeX with Postscript figures.
Theoretical Evolutionary Genetics
a draft text by
I am gradually finishing it, year by year. In the meantime:
This book is Copyright, 1978, 1983, 1988, 1991, 1992, 1994, 1995, 1997, 1999,
2001, 2003, 2005, 2007, 2009, 2011, 2013, 2015, 2016 by Joseph Felsenstein. All rights reserved.
You may download and print a copy for your own use only. You may not resell
it, transfer digital copies to anyone else, or post it anywhere for electronic
Any reprinting or resale must be negotiated with me and have my prior approval.
Click here to download a PDF (3.58Mb)
Here is a list of known errors in the 2013 edition of the book.
Notices added in compliance with University of Washington
requirements for web sites hosted at the University: